Paragnathia formica (Hesse, 1864)


P. formica is similar to Gnathia species, having only five pairs of legs and exhibiting marked sexual dimorphism. Males have large heads and mandibles and females are inflated with fused pereon somites 3-5 and reduced mouthparts. The young (praniza larvae) have a similar shape to the females. P. formica are between 2.5 and 5.0 mm in length. Unlike species in the genus Gnathia, the male pylopod (mouthpart deriving from the modified first pereopod) has five articles (in Gnathia it has two or three).

Distribution and Habitat

Young praniza larvae are external parasites of fish. Late praniza larvae and adults inhabit mud cliffs around MHWN in estuaries and saltmarshes. Adults live in colonies in short galleries, whilst praniza larvae may be scattered more generally in the habitat. Males, which live more than two years, can be found all year round, whilst females, which usually live for about a year, are mostly found between May and October. For more details on finding this species, see here. P. formica has mostly been recorded in south-western Britain, though it has also been found in Northumberland and south-west Scotland, so it is rather more widespread than the NBN map indicates.



Naylor, E. & A. Brandt. 2015. Intertidal Marine Isopods. Synopses of the British Fauna (New Series), No. 3. Field Studies Council, for The Linnean Society of London.